Dr Caroline Howlett

Essex Pathways
Dr Caroline Howlett



  • BSc (Hons) Zoology University of Leeds, (2011)

  • MScR Biological Anthropology Durham University, (2012)

  • PGCHE University of Kent, (2018)

  • PhD Anthropology University of Kent, (2019)


Other academic

  • Lecturer, Essex Pathways, University of Essex (1/8/2019 - present)

Teaching and supervision

Current teaching responsibilities

  • Introduction to Biology (IA102)


Journal articles (6)

Howlett, C. and Wheeler, BC., (2021). Prenatal Androgen Effects as a Proximate Mechanism Underpinning Variation in Social Behavior Among Female Nonhuman Primates. International Journal of Primatology. 42 (2), 301-329

Ayers, AM., Allan, ATL., Howlett, C., Tordiffe, ASW., Williams, KS., Williams, ST. and Hill, RA., (2020). Illuminating movement? Nocturnal activity patterns in chacma baboons. Journal of Zoology. 310 (4), 287-297

PARKER, EJ., HILL, RA., ALLAN, ATL., HOWLETT, C. and KOYAMA, NF., (2020). Influence of food availability, plant productivity, and indigenous forest use on ranging behavior of the endangered samango monkey ( Cercopithecus albogularis schwarzi ), in the Soutpansberg Mountains, South Africa. Integrative Zoology. 15 (5), 385-400

Howlett, C. and Hill, RA., (2017). Can zoo enclosures inform exclosure design for crop-raiding primates? A preliminary assessment. African Journal of Ecology. 55 (4), 727-730

Howlett, C., Setchell, JM., Hill, RA. and Barton, RA., (2015). The 2D:4D digit ratio and social behaviour in wild female chacma baboons (Papio ursinus) in relation to dominance, aggression, interest in infants, affiliation and heritability. Behavioral Ecology and Sociobiology. 69 (1), 61-74

Howlett, C., Marshall, AR. and Hughes, WOH., (2012). Digit Ratios and Dominance in Female Baboons (Papio hamadryas and Papio ursinus). International Journal of Primatology. 33 (6), 1439-1452

+44 (0) 1206 872231


4.111, Colchester Campus